Phyllonorycter rajella (Linnaeus, 1758)

• Overview
• Types
• DNA
• Distribution
• Plants
• Parasites
• Photos
• Children
• Synonyms

Permalink:	PHYLRAJE
Higher taxa:	Gracillariidae (family)/  Lithocolletinae (subfamily)/  Phyllonorycter (genus)/
Status:	Accepted
Authorship:	(Linnaeus, 1758)
Citation:	P.[halaena] Tinea Rajella.
Original genus:	Tinea
Publication:	Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum Classes, Ordines, Genera, Species, cum characteribus, differentiis, synonymis, Iocis. Tomus 1. Editio Decima, Reformata.
Journal:	—
Volume:
Issue:
Pagination:	542, nr. 298

Phyllonorycter rajella (1)

---

Scientific name	TypeStatus	Gender	Museum	Description	Country	Locality	Publication	URL
Phyllonorycter rajella (Linnaeus, 1758)		unknown		Not stated. Probably, Linnaeus did not see this species in natura, but based his description on the figure by de Geer (1752: pl. 31, figs. 11–12		Not stated.
Phyllonorycter rajella (Linnaeus, 1758)		unknown		see also Robinson & Schmidt Nielsen 1983: 226).		Not stated.

Distribution note:

Recorded from the U.S.A. as L.[ithocolletis] alniella Z.? (Frey & Boll 1873: 210), misidentification (see also Walsingham 1889b: 77).

Scientific name	Country	Province	Publication	URL
Phyllonorycter rajella (Linnaeus, 1758)	Albania		Rebel, H. & Zerny, H. 1931. Wissenschaftliche ergebnisse der im auftrage und mit kosten der akademie der wissenschaften in Wien im jahre 1918 entsendeten expedition nach Nordalbanien. page 156 (as alniella)
Phyllonorycter rajella (Linnaeus, 1758)	Austria		Denis, M. & Schiffermüller, I. 1775. Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend herausgegeben von einigen Lehrern am k. k. Theresianum. page 143
Phyllonorycter rajella (Linnaeus, 1758)	Belarus		Merzheerskaya, O. I., Litvinova, A. N. & Molchanova, R. V. 1976. Lepidoptera of Belorussia. Catalogue. page 17
Phyllonorycter rajella (Linnaeus, 1758)	Belgium		De Fré, C. 1858. Catalogue des Microlépidoptères de la Belgique. page 155
Phyllonorycter rajella (Linnaeus, 1758)	Bosnia and Herzegovina		Skala, H. 1937b. Einiges über Falter-Minen aus dem Mediterrangebiet. page 110 (as alniella)
Phyllonorycter rajella (Linnaeus, 1758)	Bulgaria		Buszko, J. & Beshkov, S. 2004. A preliminary survey of leafmining moths (Insecta: Lepidopetra: Microlepidoptera) of the Bulgarian part of Eastern Rhodopes. In: Beron, P. & Popov, A. (Eds.), Biodiversity of Bulgaria. 2. Biodiversity of Eastern Rhodopes (Bulgaria and Greece). page 730
Phyllonorycter rajella (Linnaeus, 1758)	Croatia		Mann, J. 1867a. Schmetterlinge gesammelt im J. 1866 um Josefsthal in der croat. Militärgrenze. page 74
Phyllonorycter rajella (Linnaeus, 1758)	Czech Republic		Nickerl, O. 1894. Catalogus insectorum faunae bohemicae III. Die Kleinschmetterlinge (Microlepidoptera) Böhmens. page 29 (as alniella)
Phyllonorycter rajella (Linnaeus, 1758)	Denmark		Larsen, C. S. 1916. Fortegnelse over Danmarks Microlepidoptera. page 217 (as alniella)
Phyllonorycter rajella (Linnaeus, 1758)	Estonia		Nolcken, J. H. W. 1870. Lepidopterologische Fauna von Estland, Livland und Kurland. Microlepidoptera. Heft. IV. page 727
Phyllonorycter rajella (Linnaeus, 1758)	Finland		Tengström, J. M. J. 1848. Bidrag till Finlands Fjäril-Fauna. page 153 (as alniella)
Phyllonorycter rajella (Linnaeus, 1758)	France		de Villers, C. J. 1789. Caroli Linnaei entomologia, Fauna Suecicae descriptionibus aucta; D. D. Scopoli, Geoffroy, De Geer, Fabricii, Schrank etc. Tomus secundus. page 493
Phyllonorycter rajella (Linnaeus, 1758)	Germany		Hübner, J. 1796–1838. Sammlung europäischer Schmetterlinge. Achte Horde. Tineae Die Schaben; nach der Natur geordnet, beschrieben und vorgestellt. page 57
Phyllonorycter rajella (Linnaeus, 1758)	Hungary		Gozmány, L. A. 1954. The examination of Microlepidoptera coenoses mining on trees. page 63
Phyllonorycter rajella (Linnaeus, 1758)	Ireland		Meyrick, E. 1927a. A revised handbook of British Lepidoptera. page 772
Phyllonorycter rajella (Linnaeus, 1758)	Italy	Sardinia	Gaedike, R., Hättenschwiler, P., Triberti, P. & Zangheri, S. 1995. 81. Lepidoptera Tineoidea I. In: Minelli, A., Ruffo, S & La Posta, S. (eds.): Checklist delle species della fauna Italiana. page 18
Phyllonorycter rajella (Linnaeus, 1758)	Italy		Zeller, P. C. 1850b. Verzeichniss der von Herrn Jos. Mann beobachteten Toscanischen Microlepidoptera. page 209 (as alniella)
Phyllonorycter rajella (Linnaeus, 1758)	Latvia		Lienig, F. 1846. Lepidopterische Fauna von Lievland und Curland. page 300
Phyllonorycter rajella (Linnaeus, 1758)	Lithuania		Ivinskis, P. & Pakalniškis, S. 1984. Microlepidoptera of Lithuania (Gracillariidae). page 29
Phyllonorycter rajella (Linnaeus, 1758)	Luxembourg		Wagner-Rollinger, C. 1974. Les lépidoptères du Grand-Duché de Luxembourg (et des régions limitrophes). page 303 (as alnifoliella)
Phyllonorycter rajella (Linnaeus, 1758)	Moldova, Republic of		Kuznetzov, V. I. & Baryshnikova, S. V. 1998. Brief catalogue of the mining moths of the fam. Gracillariidae (Lepidoptera) of Russia and adjacent countries. page 44
Phyllonorycter rajella (Linnaeus, 1758)	Montenegro		Triberti, P. & Braggio, S. 2011. Remarks on some families of leaf-mining Microlepidoptera from central-southern Sardinia, with some ecological considerations (Lepidoptera: Nepticulidae, Bucculatricidae, Gracillariidae). page 773
Phyllonorycter rajella (Linnaeus, 1758)	Netherlands		de Graaf, H. W. & Snellen, P. C. T. 1866. Microlepidoptera in Nederland waargenomen. — In: Herklots, J. A. Bouwstoffen voor eene Fauna van Nederland 3. page 290
Phyllonorycter rajella (Linnaeus, 1758)	Norway		Haanshus, K. 1933. Fortegnelse over Norges Lepidoptera. page 206 (as alniella)
Phyllonorycter rajella (Linnaeus, 1758)	Poland		Zeller, P. C. 1839. Versuch einer naturgemässen Eintheilung der Schaben. page 217
Phyllonorycter rajella (Linnaeus, 1758)	Romania		Caradja, A. 1899. Zusammenstellung der bisher in Rumänien beobachteten Microlepidopteren. page 213 (as alniella)
Phyllonorycter rajella (Linnaeus, 1758)	Russian Federation	European Part	Wocke, M. F. 1871. II. Microlepidoptera. In Staudinger, O. & Wocke, M. F. Catalog der Lepidopteren des europaeischen Faunengebiets. page 329
Phyllonorycter rajella (Linnaeus, 1758)	Serbia		Triberti, P. & Braggio, S. 2011. Remarks on some families of leaf-mining Microlepidoptera from central-southern Sardinia, with some ecological considerations (Lepidoptera: Nepticulidae, Bucculatricidae, Gracillariidae). page 773
Phyllonorycter rajella (Linnaeus, 1758)	Slovakia		Skala, H. 1937a. Minen aus Mittel- und Südeuropa (Fortsetzung und Schluss). page 19 (as alniella)
Phyllonorycter rajella (Linnaeus, 1758)	Slovenia		Gutzwiller, F., Dedeine, F., Kaiser, W., Giron, D. & Lopez-Vaamonde, C. 2015. Correlation between the green-island phenotype and Wolbachia infections during the evolutionary diversification of Gracillariidae leaf-mining moths. page suppl. table S1
Phyllonorycter rajella (Linnaeus, 1758)	Sweden		Linnaeus, C. 1761. Fauna Svecica, sistens Animalia Sveciae Regni: Mammalia, Aves, Amphibia, Insecta, Vermes. Distributa per classes & ordines, genera & species, cum differentiis specierum, synonymis auctorum, nominibus incolarum, locis natalium, descriptionibus Insectorum. page 362
Phyllonorycter rajella (Linnaeus, 1758)	Switzerland		Frey, H. 1855. Ueber die in der Schweiz beobachteten Arten des Genus Lithocolletis. Zell. page 622 (as alniella)
Phyllonorycter rajella (Linnaeus, 1758)	Ukraine		Kuznetzov, V. I. & Baryshnikova, S. V. 1998. Brief catalogue of the mining moths of the fam. Gracillariidae (Lepidoptera) of Russia and adjacent countries. page 44
Phyllonorycter rajella (Linnaeus, 1758)	United Kingdom		Haworth, A. H. 1828. Lepidopterae Britannicae. Pars IV cum indice finali. page 577

Plant note:

Blotch mine on underside of leaf.

Scientific name	Plant	Plant Family	Country	Publication	URL
Phyllonorycter rajella (Linnaeus, 1758)	Alnus alnobetula	Betulaceae	Italy	Hartig, F. 1964. Microlepidotteri della Venezia Tridentina e delle regioni adiacenti. Parte III. (Fam. Gelechiidae - Micropterigidae). page 178
Phyllonorycter rajella (Linnaeus, 1758)	Alnus alnobetula	Betulaceae	Austria	Skala, H. 1951. Minen aus Steiermark, gesammelt von Fritz Hoffmann. page 119
Phyllonorycter rajella (Linnaeus, 1758)	Alnus cordata	Betulaceae	Denmark	Larsen, C. S. 1916. Fortegnelse over Danmarks Microlepidoptera. page 217 (as alniella)
Phyllonorycter rajella (Linnaeus, 1758)	Alnus cordata	Betulaceae	United Kingdom	Emmet, A. M. 1988. A field guide to the smaller British Lepidoptera. Second edition. page 65
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	Bosnia and Herzegovina	Skala, H. 1937b. Einiges über Falter-Minen aus dem Mediterrangebiet. page 110
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	Poland	Brischke, C. G. A. 1880. Die Blattminirer in Danzig's Umgebung. page 51
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	Slovakia	Hrubý, K. 1964. Prodromus Lepidopterorum Slovaciae. page 200
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	Switzerland	Frey, H. 1855. Ueber die in der Schweiz beobachteten Arten des Genus Lithocolletis. Zell. page 622
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	Czech Republic	Gregor, F. & Povolný, D. 1950b. The members of Lithocolletis Hb. mining Acer and Alnus. page 130
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	Italy	Mendes D'Azevedo, C. 1910b. Lepidopteros Romanos (Roma, Castelli Romani). page 148
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	Austria	Mitterberger, K. 1909. Verzeichnis der im Kronlande Salzburg bisher beobachteten Mikrolepidopteren. page 291
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	France	Le Marchand, S. 1936. Clé ou table analytique pour la détermination des espèces françaises de Lithocolletis (Famille des Gracilariidae [sic]). page 93
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	United Kingdom	Meyrick, E. 1927a. A revised handbook of British Lepidoptera. page 772
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	Belgium	De Fré, C. 1858. Catalogue des Microlépidoptères de la Belgique. page 156
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	Sweden	Wallengren, H. J. D. 1883. Skandinaviens arter af tineidgruppen Lithocolletidae (Staint). page 199
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	Netherlands	de Graaf, H. W. & Snellen, P. C. T. 1866. Microlepidoptera in Nederland waargenomen. — In: Herklots, J. A. Bouwstoffen voor eene Fauna van Nederland 3. page 290
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	Estonia	Petersen, W. 1930. Nachträge zur Lepidopterenfauna von Estland, mit besonderer Berüksichtigung der Gattungen Lithocolletis und Nepticula. page 90
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	Denmark	Larsen, C. S. 1916. Fortegnelse over Danmarks Microlepidoptera. page 217 (as alniella)
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	Lithuania	Ivinskis, P. & Pakalniškis, S. 1984. Microlepidoptera of Lithuania (Gracillariidae). page 29
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	Croatia	Matoševic, D., Pernek, M., Dubravac, T. & Baric, B. 2009. Research of leafminers on woody plants in Croatia. page 385
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	Slovenia	Gutzwiller, F., Dedeine, F., Kaiser, W., Giron, D. & Lopez-Vaamonde, C. 2015. Correlation between the green-island phenotype and Wolbachia infections during the evolutionary diversification of Gracillariidae leaf-mining moths. page suppl. table S1
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	Albania	Gutzwiller, F., Dedeine, F., Kaiser, W., Giron, D. & Lopez-Vaamonde, C. 2015. Correlation between the green-island phenotype and Wolbachia infections during the evolutionary diversification of Gracillariidae leaf-mining moths. page suppl. table S1
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	Bulgaria	Buszko, J. & Beshkov, S. 2004. A preliminary survey of leafmining moths (Insecta: Lepidopetra: Microlepidoptera) of the Bulgarian part of Eastern Rhodopes. In: Beron, P. & Popov, A. (Eds.), Biodiversity of Bulgaria. 2. Biodiversity of Eastern Rhodopes (Bulgaria and Greece). page 730
Phyllonorycter rajella (Linnaeus, 1758)	Alnus glutinosa	Betulaceae	Germany	Hübner, J. 1796–1838. Sammlung europäischer Schmetterlinge. Achte Horde. Tineae Die Schaben; nach der Natur geordnet, beschrieben und vorgestellt. page 57
Phyllonorycter rajella (Linnaeus, 1758)	Alnus incana	Betulaceae	Sweden	Wallengren, H. J. D. 1883. Skandinaviens arter af tineidgruppen Lithocolletidae (Staint). page 199
Phyllonorycter rajella (Linnaeus, 1758)	Alnus incana	Betulaceae	Lithuania	Puplesiene, J. & Noreika, R. 1993. A brief karyological review of the Gracillariidae (Lepidoptera). page 57
Phyllonorycter rajella (Linnaeus, 1758)	Alnus incana	Betulaceae	Italy	Klimesch, J. 1950. Contibuto alla fauna Lepidopterologica del Trentino. page 48
Phyllonorycter rajella (Linnaeus, 1758)	Alnus incana	Betulaceae	Germany	Steudel, W. & Hofmann, E. 1882. Verzeichniss württembergischer Kleinschmetterlinge. page 230
Phyllonorycter rajella (Linnaeus, 1758)	Alnus incana	Betulaceae	Austria	Galvagni, E. 1955. Schmetterlinge aus dem westlichen Alpengebiet Niederösterreichs aus Waidhofen an der Ybbs, Ybbsitz, Lunz und dem Ybbstal, dem Minichholt bei Steyr (N.-Ö.) und Umgebung; aus Türnitz, dem Ötschergebiet, Annaberg und der Walster (Prodromus Zone 1 und 4). page 173
Phyllonorycter rajella (Linnaeus, 1758)	Alnus incana	Betulaceae	Czech Republic	Gregor, F. & Povolný, D. 1950b. The members of Lithocolletis Hb. mining Acer and Alnus. page 130
Phyllonorycter rajella (Linnaeus, 1758)	Alnus japonica	Betulaceae	Germany	Osthelder, L. 1951. Die Schmetterlinge Südbayerns und der angrenzenden nördlichen Kalkalpen. II. Teil. Die Kleischmetterlinge. 2. Heft. Glyphipterygidae bis Micropterygidae. page 218
Phyllonorycter rajella (Linnaeus, 1758)	Alnus sp.	Betulaceae	Lithuania	Ivinskis, P., Pakalniškis, S. & Puplesis, R. 1985. Augalus minuojantys vabzdziai. page 95
Phyllonorycter rajella (Linnaeus, 1758)	Alnus sp.	Betulaceae	Germany	Eckstein, K. 1933. Die Schmetterlinge Deutschlands mit besonderer Berücksichtigung ihrer Biologie und wirtschaftlichen Bedeutung. 5. Band Die Kleinschmetterlinge Deutschlands. page 160
Phyllonorycter rajella (Linnaeus, 1758)	Alnus sp.	Betulaceae	Latvia	Lienig, F. 1846. Lepidopterische Fauna von Lievland und Curland. page 300
Phyllonorycter rajella (Linnaeus, 1758)	Alnus sp.	Betulaceae	Sweden	Benander, P. 1944. Sveriges Lithocolletider (Gracilariidae) [sic]. page 100
Phyllonorycter rajella (Linnaeus, 1758)	Alnus sp.	Betulaceae		Fabricius, J. C. 1781. Species insectorum exhibentes eorum differentias specificas, synonyma autorum, loca natalia, metamorphosin adiectis observationibus, descriptionibus. Tom II. page 304
Phyllonorycter rajella (Linnaeus, 1758)	Alnus sp.	Betulaceae	Estonia	Kesküla, T., Martin, M., Pedmanson, R. & Miller, R. 1996. Records of some Microlepidoptera in Estonia. page 11
Phyllonorycter rajella (Linnaeus, 1758)	Alnus sp.	Betulaceae	United Kingdom	Emmet, A. M. 1988. A field guide to the smaller British Lepidoptera. Second edition. page 65
Phyllonorycter rajella (Linnaeus, 1758)	Alnus viridis subsp. sinuata	Betulaceae	Germany	Osthelder, L. 1951. Die Schmetterlinge Südbayerns und der angrenzenden nördlichen Kalkalpen. II. Teil. Die Kleischmetterlinge. 2. Heft. Glyphipterygidae bis Micropterygidae. page 218

Scientific name	Parasitoid	Parasitoid Family	Country	Publication	URL
Phyllonorycter rajella (Linnaeus, 1758)	Achrysocharoides splendens	Eulophidae	Germany	Niehus, O. 1986. Zur Biologie und Ökologie einiger Erlen-Blattütenmotten (Phyllonorycter Hübner; Lep., Gracilariidae) und ihrer Parasiten. page 72
Phyllonorycter rajella (Linnaeus, 1758)	Achrysocharoides splendens	Eulophidae	United Kingdom	Rott, A. S. & Godfray, H. C. J. 2000. The structure of leafminer-parasitoid community. page 278
Phyllonorycter rajella (Linnaeus, 1758)	Achrysocharoides splendens	Eulophidae	Poland	Vidal, S. & Buszko, J. 1990. Studies on the mining Lepidoptera of Poland. VIII. Chalcidoid wasps reared from mining Lepidoptera (Hymenoptera, Chalcidoidea). page 77
Phyllonorycter rajella (Linnaeus, 1758)	Agathis pedias	Braconidae	Germany	Niehus, O. 1986. Zur Biologie und Ökologie einiger Erlen-Blattütenmotten (Phyllonorycter Hübner; Lep., Gracilariidae) und ihrer Parasiten. page 72
Phyllonorycter rajella (Linnaeus, 1758)	Ageniaspis nepticulae	Encyrtidae	United Kingdom	Askew, R. R. 1983. A revision of the encyrtid genus Holcothorax (Hymenoptera). page 135
Phyllonorycter rajella (Linnaeus, 1758)	Ageniaspis testaceipes	Encyrtidae	Belgium	De Prins, W. 1983. Parasitaire Hymenoptera gekweekt uit mijnen van Phyllonorycter soorten (Hymenoptera; Lepidoptera). page 68
Phyllonorycter rajella (Linnaeus, 1758)	Ageniaspis testaceipes	Encyrtidae	Germany	Niehus, O. 1986. Zur Biologie und Ökologie einiger Erlen-Blattütenmotten (Phyllonorycter Hübner; Lep., Gracilariidae) und ihrer Parasiten. page 72
Phyllonorycter rajella (Linnaeus, 1758)	Apanteles bicolor	Braconidae	Poland	Marczak, P. & Buszko, J. 1993. Braconid wasps (Hypneoptera, Braconidae) reared from mining Lepidoptera. page 268
Phyllonorycter rajella (Linnaeus, 1758)	Apanteles nanus	Braconidae	Germany	Niehus, O. 1986. Zur Biologie und Ökologie einiger Erlen-Blattütenmotten (Phyllonorycter Hübner; Lep., Gracilariidae) und ihrer Parasiten. page 72
Phyllonorycter rajella (Linnaeus, 1758)	Apanteles sp.	Braconidae	United Kingdom	Rott, A. S. & Godfray, H. C. J. 2000. The structure of leafminer-parasitoid community. page 278
Phyllonorycter rajella (Linnaeus, 1758)	Chrysocharis laomedon	Eulophidae	Germany	Niehus, O. 1986. Zur Biologie und Ökologie einiger Erlen-Blattütenmotten (Phyllonorycter Hübner; Lep., Gracilariidae) und ihrer Parasiten. page 72
Phyllonorycter rajella (Linnaeus, 1758)	Chrysocharis laomedon	Eulophidae	Poland	Vidal, S. & Buszko, J. 1990. Studies on the mining Lepidoptera of Poland. VIII. Chalcidoid wasps reared from mining Lepidoptera (Hymenoptera, Chalcidoidea). page 80
Phyllonorycter rajella (Linnaeus, 1758)	Chrysocharis laomedon	Eulophidae	United Kingdom	Rott, A. S. & Godfray, H. C. J. 2000. The structure of leafminer-parasitoid community. page 278
Phyllonorycter rajella (Linnaeus, 1758)	Chrysocharis nephereus	Eulophidae	Germany	Niehus, O. 1986. Zur Biologie und Ökologie einiger Erlen-Blattütenmotten (Phyllonorycter Hübner; Lep., Gracilariidae) und ihrer Parasiten. page 72
Phyllonorycter rajella (Linnaeus, 1758)	Chrysocharis nephereus	Eulophidae	Belgium	De Prins, W. 1983. Parasitaire Hymenoptera gekweekt uit mijnen van Phyllonorycter soorten (Hymenoptera; Lepidoptera). page 70
Phyllonorycter rajella (Linnaeus, 1758)	Chrysocharis nitetis	Eulophidae	Belgium	De Prins, W. 1983. Parasitaire Hymenoptera gekweekt uit mijnen van Phyllonorycter soorten (Hymenoptera; Lepidoptera). page 70
Phyllonorycter rajella (Linnaeus, 1758)	Cirrospilus diallus	Eulophidae	United Kingdom	Rott, A. S. & Godfray, H. C. J. 2000. The structure of leafminer-parasitoid community. page 278
Phyllonorycter rajella (Linnaeus, 1758)	Cirrospilus diallus	Eulophidae	Poland	Górny, S. 1979. Parasitic Hymenoptera on black alder, Alnus glutinosa (L.) Gaertn., near Ostróda, Poland. page 345
Phyllonorycter rajella (Linnaeus, 1758)	Cirrospilus elegantissimus	Eulophidae	Germany	Niehus, O. 1986. Zur Biologie und Ökologie einiger Erlen-Blattütenmotten (Phyllonorycter Hübner; Lep., Gracilariidae) und ihrer Parasiten. page 72
Phyllonorycter rajella (Linnaeus, 1758)	Cirrospilus lyncus	Eulophidae	United Kingdom	Rott, A. S. & Godfray, H. C. J. 2000. The structure of leafminer-parasitoid community. page 278
Phyllonorycter rajella (Linnaeus, 1758)	Cirrospilus pictus	Eulophidae	Germany	Niehus, O. 1986. Zur Biologie und Ökologie einiger Erlen-Blattütenmotten (Phyllonorycter Hübner; Lep., Gracilariidae) und ihrer Parasiten. page 72
Phyllonorycter rajella (Linnaeus, 1758)	Cirrospilus viticola	Eulophidae	Poland	Górny, S. 1979. Parasitic Hymenoptera on black alder, Alnus glutinosa (L.) Gaertn., near Ostróda, Poland. page 346
Phyllonorycter rajella (Linnaeus, 1758)	Cirrospilus vittatus	Eulophidae	Germany	Niehus, O. 1986. Zur Biologie und Ökologie einiger Erlen-Blattütenmotten (Phyllonorycter Hübner; Lep., Gracilariidae) und ihrer Parasiten. page 72
Phyllonorycter rajella (Linnaeus, 1758)	Colastes braconius	Braconidae	Germany	Niehus, O. 1986. Zur Biologie und Ökologie einiger Erlen-Blattütenmotten (Phyllonorycter Hübner; Lep., Gracilariidae) und ihrer Parasiten. page 72
Phyllonorycter rajella (Linnaeus, 1758)	Colastes braconius	Braconidae	United Kingdom	Shaw, M. R. & Askew, R. R. 1976. Ichneumonoidea (Hymenoptera) parasitic upon leaf-mining insects of the orders Lepidoptera, Hymenoptera and Coleoptera. page 128
Phyllonorycter rajella (Linnaeus, 1758)	Itoplectis alternans	Ichneumonidae	Poland	Górny, S. 1979. Parasitic Hymenoptera on black alder, Alnus glutinosa (L.) Gaertn., near Ostróda, Poland. page 309
Phyllonorycter rajella (Linnaeus, 1758)	Minotetrastichus frontalis	Eulophidae	Germany	Niehus, O. 1986. Zur Biologie und Ökologie einiger Erlen-Blattütenmotten (Phyllonorycter Hübner; Lep., Gracilariidae) und ihrer Parasiten. page 72
Phyllonorycter rajella (Linnaeus, 1758)	Pediobius alcaeus	Eulophidae	Netherlands	Shaw, M. R. & Askew, R. R. 1999. Reproductive variability in Pediobius alcaeus (Walker) (Hymenoptera: Eulophidae), a parasitoid of Phyllonorycter (Lepidoptera: Gracillariidae). page 128
Phyllonorycter rajella (Linnaeus, 1758)	Pediobius alcaeus	Eulophidae	Germany	Niehus, O. 1986. Zur Biologie und Ökologie einiger Erlen-Blattütenmotten (Phyllonorycter Hübner; Lep., Gracilariidae) und ihrer Parasiten. page 72
Phyllonorycter rajella (Linnaeus, 1758)	Pediobius alcaeus	Eulophidae	United Kingdom	Shaw, M. R. & Askew, R. R. 1999. Reproductive variability in Pediobius alcaeus (Walker) (Hymenoptera: Eulophidae), a parasitoid of Phyllonorycter (Lepidoptera: Gracillariidae). page 128
Phyllonorycter rajella (Linnaeus, 1758)	Pediobius alcaeus	Eulophidae	Belgium	De Prins, W. 1983. Parasitaire Hymenoptera gekweekt uit mijnen van Phyllonorycter soorten (Hymenoptera; Lepidoptera). page 70
Phyllonorycter rajella (Linnaeus, 1758)	Pediobius saulius	Eulophidae	United Kingdom	Rott, A. S. & Godfray, H. C. J. 2000. The structure of leafminer-parasitoid community. page 278
Phyllonorycter rajella (Linnaeus, 1758)	Pediobius saulius	Eulophidae	Belgium	De Prins, W. 1983. Parasitaire Hymenoptera gekweekt uit mijnen van Phyllonorycter soorten (Hymenoptera; Lepidoptera). page 70
Phyllonorycter rajella (Linnaeus, 1758)	Pediobius saulius	Eulophidae	Germany	Niehus, O. 1986. Zur Biologie und Ökologie einiger Erlen-Blattütenmotten (Phyllonorycter Hübner; Lep., Gracilariidae) und ihrer Parasiten. page 72
Phyllonorycter rajella (Linnaeus, 1758)	Pimpla turionellae	Ichneumonidae	Poland	Górny, S. 1979. Parasitic Hymenoptera on black alder, Alnus glutinosa (L.) Gaertn., near Ostróda, Poland. page 311
Phyllonorycter rajella (Linnaeus, 1758)	Pnigalio longulus	Eulophidae	Germany	Niehus, O. 1986. Zur Biologie und Ökologie einiger Erlen-Blattütenmotten (Phyllonorycter Hübner; Lep., Gracilariidae) und ihrer Parasiten. page 72
Phyllonorycter rajella (Linnaeus, 1758)	Pnigalio pectinicornis	Eulophidae	Poland	Vidal, S. & Buszko, J. 1990. Studies on the mining Lepidoptera of Poland. VIII. Chalcidoid wasps reared from mining Lepidoptera (Hymenoptera, Chalcidoidea). page 93
Phyllonorycter rajella (Linnaeus, 1758)	Pnigalio pectinicornis	Eulophidae	Germany	Niehus, O. 1986. Zur Biologie und Ökologie einiger Erlen-Blattütenmotten (Phyllonorycter Hübner; Lep., Gracilariidae) und ihrer Parasiten. page 72
Phyllonorycter rajella (Linnaeus, 1758)	Pnigalio soemius	Eulophidae	Germany	Niehus, O. 1986. Zur Biologie und Ökologie einiger Erlen-Blattütenmotten (Phyllonorycter Hübner; Lep., Gracilariidae) und ihrer Parasiten. page 72
Phyllonorycter rajella (Linnaeus, 1758)	Scambus calobatus	Ichneumonidae	Poland	Górny, S. 1979. Parasitic Hymenoptera on black alder, Alnus glutinosa (L.) Gaertn., near Ostróda, Poland. page 307
Phyllonorycter rajella (Linnaeus, 1758)	Sympiesis gordius	Eulophidae	Germany	Niehus, O. 1986. Zur Biologie und Ökologie einiger Erlen-Blattütenmotten (Phyllonorycter Hübner; Lep., Gracilariidae) und ihrer Parasiten. page 72
Phyllonorycter rajella (Linnaeus, 1758)	Sympiesis gordius	Eulophidae	United Kingdom	Rott, A. S. & Godfray, H. C. J. 2000. The structure of leafminer-parasitoid community. page 278
Phyllonorycter rajella (Linnaeus, 1758)	Sympiesis gordius	Eulophidae	Poland	Górny, S. 1979. Parasitic Hymenoptera on black alder, Alnus glutinosa (L.) Gaertn., near Ostróda, Poland. page 347
Phyllonorycter rajella (Linnaeus, 1758)	Sympiesis grahami	Eulophidae	United Kingdom	Rott, A. S. & Godfray, H. C. J. 2000. The structure of leafminer-parasitoid community. page 278
Phyllonorycter rajella (Linnaeus, 1758)	Sympiesis sericeicornis	Eulophidae	United Kingdom	Rott, A. S. & Godfray, H. C. J. 2000. The structure of leafminer-parasitoid community. page 278
Phyllonorycter rajella (Linnaeus, 1758)	Sympiesis sericeicornis	Eulophidae	Germany	Niehus, O. 1986. Zur Biologie und Ökologie einiger Erlen-Blattütenmotten (Phyllonorycter Hübner; Lep., Gracilariidae) und ihrer Parasiten. page 72
Phyllonorycter rajella (Linnaeus, 1758)	Sympiesis sericeicornis	Eulophidae	Poland	Górny, S. 1979. Parasitic Hymenoptera on black alder, Alnus glutinosa (L.) Gaertn., near Ostróda, Poland. page 347

Phyllonorycter rajella (1)

Phyllonorycter rajella (2)

Phyllonorycter rajella (3)

Phyllonorycter alniella (1)

Phyllonorycter alnifoliella (1)

Scientific Name	Authorship	Rank	Status	Original Combination

Scientific Name	Authorship	Rank	Status	Original Combination
Phyllonorycter alniella (Zeller, 1846)	(Zeller, 1846)	species	Synonym	Lithocolletis alniella
Phyllonorycter alnifoliella (Hübner, 1796)	(Hübner, 1796)	species	Synonym	Tinea alnifoliella
Phyllonorycter neapolitana (Rebel, 1901)	(Rebel, 1901)	species	Synonym	Lithocolletis neapolitana
Phyllonorycter salincolella (Sorhagen, 1900)	(Sorhagen, 1900)	species	Synonym	Lithocolletis salincolella